Almodóvar, Ana; Nicola, Graciela G. & Elvira, Benigno (2008): Natural hybridization of Barbus bocagei x Barbus comizo (Cyprinidae) in Tagus River basin, central Spain [English with French abstract]. Cybium32(2): 99–102. PDF fulltext
de Graaf, Martin; Megens, Hendrik-Jan; Samallo, Johannis & Sibbing, Ferdinand A. (2007): Evolutionary origin of Lake Tana's (Ethiopia) small Barbus species: indications of rapid ecological divergence and speciation. Anim. Biol.57(1): 39–48. doi:10.1163/157075607780002069 (HTML abstract)